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The goal of this 11th New Phytologist Symposium is to highlight areas of plant speciation where considerable progress has been made or is likely in the near future and to relate these discoveries to the work of early students of plant speciation, particularly Verne Grant. Perhaps no other scientist has contributed more to our understanding of plant speciation or influenced more plant evolutionists, young and old. To honour his many empirical discoveries, conceptual advances, and masterful syntheses of the speciation literature, the symposium is commemoratively named after his most famous book, Plant Speciation.