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center dot Global warming and droughts push forests closer to their thermal limits, altering tree carbon uptake and growth. To prevent critical overheating, trees can adjust their thermotolerance (T-crit), temperature and photosynthetic optima (T-opt and A(opt)), and canopy temperature (T-can) to stay below damaging thresholds. However, we lack an understanding of how soil droughts affect photosynthetic thermal plasticity and T-can regulation.